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Recent genetic data show that ground hornbills and Bycanistes form a clade outside the rest of the hornbill lineage.[5] They are thought to represent an early African lineage, while the rest of Bucerotiformes evolved in Asia. The hoopoe subspecies Saint Helena hoopoe and the Madagascar subspecies are sometimes elevated to a full species. The two wood hoopoe genera, Phoeniculus and Rhinopomastus, appear to have diverged about 10 million years ago, so some systematists treat them as separate subfamilies or even separate families.[6]
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